How were the bolts installed at the factory?

And I'm confused that the holes they're going into are round, not square. What keeps/kept the bolt from spinning? Am I misunderstanding something?

1950 Chevrolet 3100 (Ol' Roy)

1939 Packard Standard Eight Coupe (The Phantom) | 1956 Cadillac Coupe de Ville (The Bismarck) | 1964 Ford Galaxie 500 coupe |1966 Ford Mustang (Little Red) | 1969 Pontiac GTO (The Dishpit) I 1979 Ford F-100 | 1976 Ford F-150 (Big Red) | 1995 Ford F-150 (Newt)