I have a round hole in the center underneath my seat; and in my case that area is sloped inwards (like a drain). When I was prepping for the floor repairs I power washed the interior and I can state with certainty that it works effectively like a drain, although why you would need one under there I'm not sure. I put a rubber plug in mine (they had the right size at NAPA).
Picture shows where it is but doesn't do the slopes justice, IIRC it's about a 1/2" hole.
1949 Chevrolet 3/4 Ton - Still Solid. Regular Driver OT Vehicles: 1925 Hudson Brougham Super Six 1950 Chevrolet Styline (Parts) 1952 Canuck Pontiac Sedan Delivery (Well Underway) 1973 F250 4x4 Highboy 1977 F250 4x4 Lowboy