The donor wheels really only had to be the correct width. Could have been any bolt pattern, and either new or used. Remember somebody counted LHD's hand holes. The flatness of the face suggests there's really no other likely explanation for the technique used to do them. If the donor wheels were new to start with, and like Grigg says probably a bolt pattern that wasn't in big demand, it's reasonable that they could be priced at $200+ retail per wheel before the custom work was even started.
I bought a set of remanufactured 19.5s (5 lug x 8") a while back from a different company that had been returned by another customer. The customer didn't think they were balanced well enough for motor home highway use, but for my slow old rigs they're fine. On these they'd remounted old centers in new tubeless outer rims. I paid what I was told were the material costs of $85 each. Assuming that's close to correct, there's a fair amount of profit beyond labor costs in any of these custom wheels. Stu