I don't know what the differences are between a 1940 radiator filler neck and a 1941 radiator (that's what's on my truck) but the 4pound radiator cap on my truck has two seals like a modern car and functions the same way.
Whether it functions as a return reservoir or not, it can still act as a recovery tank so that excess fluid doesn't dump on the ground. The radiator could be over-filled without wasting the coolant that dumps out.